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In 1957, second-grader Bryce Loski first moves in and meets Julianna “Juli” Baker. When they look into each other’s eyes, Juli knows it’s love, but Bryce isn’t so sure and tries to avoid her.
Four years later in 1961, Bryce tries to get rid of Juli by dating Sherry Stalls, whom Juli finds unreliable. However, his best friend Garrett takes an interest in Sherry and eventually tells her the truth about the advice, therefore not taking it too well. After Juli finds out they broke up, she thinks she could have him back, but they reconsider their own choices as time goes on.
Chet Duncan, Bryce’s grandfather, moves in to live with the family. At one point, Juli falls in love with a large, old sycamore tree, but her relationship doesn’t last long after the tree itself is cut down, much to her dismay. However, her father gives her a painting of the same kind of tree, causing her to overcome her distress. Juli is hurt when she finds out Bryce has been throwing away the eggs she offered, out of fear of salmonella. Chet helps Juli fix up her yard, while Bryce begins to develop feelings for Juli.
After Juli returns home from visiting her mentally-disabled uncle Daniel, she overhears Bryce supporting Garrett’s badmouthing of her, causing her to stop having any interest in him. The Loskis invite the Bakers over for dinner and once arriving there, Juli accuses Bryce about what he did and eventually rejects him. After dinner, she apologizes for her behavior. Bryce has mixed feelings afterwards, since he isn’t forgiven and she doesn’t care enough to hold a grudge.
As the basket boy auction approaches, Juli finds out Sherry is planning to bid on Bryce. Bryce worries about what will happen in reality if Juli tries to start a bidding war with Sherry, but she bids on a boy named Eddie Trulock out of sympathy as nobody was bidding for him. However, during lunch, Bryce can’t seem to ignore Juli talking with Eddie, while he talks with Sherry. Much to his jealousy, he gets up out of his table and walks towards her. He grabs her shoulders and tries to kiss her, causing her to dodge him and leave in humiliation. Afterwards, Juli’s feelings are hurt. Garrett and Bryce both end their friendship after a disagreement about Juli. Bryce tries to talk to Juli, but because of what he did, she refuses to do so.
Two days later, Bryce plants a sapling sycamore tree in Juli’s front yard to show her how he feels. When she sees this, she goes out to help him and realizes that after all these years, they have never started talking with each other. As they plant the tree, they put their hands on each other, look into each other’s eyes, and share loving smiles.